Evidence mapPaperPMID 41888955Full record

ArticleBiology of sex differences2026

Sex-specific changes in the hippocampal proteome of Negr1

Srirathi Muthuraman, Mohan Jayaram, Liisi Promet, Toomas Jagomäe, Arun Kumar Devarajan, Andreas-Christian Hade, Mari-Anne Philips, Katyayani Singh, Eero Vasar

Abstract read
In one paragraph

Article in Biology of sex differences, 2026. The graph could read no effect estimate from its abstract, so it casts no vote on the map. Not yet cited in PubMed.

0numbers the graph read from it
0cells of the map it votes in
0citing papers in PubMed
field-weighted citation impact
1 · What the graph read from it

What it found

Each row is one number read from the abstract, on the scale the paper reported it, with its interval. Left of the dashed line favours the treatment, right favours the comparator. Under each row is the sentence it came from. New to these charts? A ten-minute tutorial.

The abstract states no effect estimate the extractor could read, or names no intervention and outcome on the map, so this paper lights no cell and moves no belief. It is still indexed, cited and linked below.

2 · The registry

The trial behind it

Trials whose registry record cites this paper, or whose number appears in the abstract. A trial that started after this paper was published is citing it as background, not reporting it.

Neither the registry nor the abstract names a trial number. If this is a trial report, that itself is worth knowing.

3 · Its place in the literature

Who cites it

0 citing papers in PubMed.

No citing paper in PubMed yet.

4 · The record

Corrections and comments

PubMed lists nothing against this paper. Absence here is not a guarantee, only a check that was made.

5 · Who and what money

Authors and funding

9 authors.

Srirathi MuthuramanInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.
Mohan JayaramInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.
Liisi PrometInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.
Toomas JagomäeInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.
Arun Kumar DevarajanInstitute of Molecular and Cell Biology, University of Tartu, Tartu, Estonia.
Andreas-Christian HadeDepartment of Pathological Anatomy and Forensic Medicine, University of Tartu, 19 Ravila Street, Tartu, 50411, Estonia.
Mari-Anne PhilipsInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.
Katyayani SinghInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia. katyayani.singh@ut.ee.
Eero VasarInstitute of Biomedicine and Translational Medicine, Department of Physiology, University of Tartu, Tartu, Estonia.

Funding

Estonian Research Foundation, the Estonian Research Council PRG685 and PRG2544, TARISTU24-TK22
6 · The paper itself

Abstract

backgroundNeuronal Growth Regulator 1 (NEGR1) is a cell adhesion molecule involved in hippocampal circuit development and function. Human genetic studies have identified NEGR1 variants as risk factors for a broad spectrum of neuropsychiatric disorders. These disorders often display sex-specific differences in prevalence, progression, and behavioral impairment, reflecting underlying maladaptive changes in neural circuitry. Findings from preclinical studies using Negr1−/− mice show several hippocampal-based behavioral and anatomical endophenotypes relevant to neuropsychiatric disorders. The hippocampus, a key region implicated in these disorders, exhibits sex-dependent anatomical features that may shape the functional impact of Negr1. However, the mechanisms driving these sex-specific characteristics have not yet been elucidated. Here, we uncover sex-specific molecular signatures and pathways associated with Negr1, using Negr1−/− mice, a genetically relevant animal model for neuropsychiatric risk.

methodsWe performed label-free quantitative proteomic analysis using eight replicates of hippocampi dissected from male and female wild-type, and Negr1−/− mice. Differentially abundant proteins were subjected to functional annotation for Gene Ontology and Protein-Protein interaction using STRING analysis. NEGR1 cellular localization was examined by immunofluorescent in rat brain and human hippocampal sections.

resultsDifferential expression analysis identified 232 proteins in males and 172 in females. STRING analysis revealed sex-specific regulation of proteins. In males, proteins linked to neurofilament organization, myelin integrity, and postsynaptic structure were downregulated, with parvalbumin (Pvalb, PV) around the central node. In contrast, proteins related to mitochondrial and stress-response pathways were upregulated. Female Negr1−/− hippocampus showed downregulation of proteins involved in translation and amide biosynthetic processes. Colocalization of NEGR1 with PV interneurons in the rat brain and the human hippocampus was observed.

conclusionsWe demonstrate, for the first time, distinct sex differences in the hippocampal proteome and identify molecular networks in Negr1−/− mice. Co-localization of NEGR1 and PV in human brain tissue provides anatomical and translational validation of a proteomic target. These findings provide new insight, offering a valuable resource for understanding NEGR1-related sex-specific mechanisms in neuropsychiatric disorders.

Indexed as

Cell Adhesion Molecules, NeuronalHippocampusMental DisordersProteomeSex CharacteristicsAnimalsFemaleMaleMiceMice, Inbred C57BLMice, KnockoutCell Adhesion Molecules, NeuronalProteomeAnimal modelHippocampusNegr1Neuropsychiatric disorderProteomicsPVPvalbSex-differenceSex-specific

Identifiers

PMID41888955
PMCPMC13141394

What Socratic holds

Textmetadata
LicenceCC BY
Read underepoch 390

Registered trials

None linked

Read under generation 80e0d062 · epoch 390. Bibliography from PubMed, PubMed Central and OpenAlex; grants from NIH RePORTER; trial links from ClinicalTrials.gov; estimates, votes and beliefs from the Socratic graph.