Evidence mapPaperPMID 42274693Full record

ArticleNanomaterials (Basel, Switzerland)2026

Cryogenic Electron Microscopy of Extracellular Vesicles from Temozolomide-Treated Glioblastoma Cells Reveals Great Morphological Heterogeneity.

Mariana Karimova, Giordana Ponziani, Diana Vardanyan, Andrea Alfieri, Maurizio Zuccotti, Stefano Tacconi, Luciana Dini

Abstract read
In one paragraph

Article in Nanomaterials (Basel, Switzerland), 2026. The graph could read no effect estimate from its abstract, so it casts no vote on the map. Cited by 1 paper.

0numbers the graph read from it
0cells of the map it votes in
1citing papers in PubMed
field-weighted citation impact
1 · What the graph read from it

What it found

Each row is one number read from the abstract, on the scale the paper reported it, with its interval. Left of the dashed line favours the treatment, right favours the comparator. Under each row is the sentence it came from. New to these charts? A ten-minute tutorial.

The abstract states no effect estimate the extractor could read, or names no intervention and outcome on the map, so this paper lights no cell and moves no belief. It is still indexed, cited and linked below.

2 · The registry

The trial behind it

Trials whose registry record cites this paper, or whose number appears in the abstract. A trial that started after this paper was published is citing it as background, not reporting it.

Neither the registry nor the abstract names a trial number. If this is a trial report, that itself is worth knowing.

3 · Its place in the literature

Who cites it

1 citing paper in PubMed.

  1. Article
4 · The record

Corrections and comments

PubMed lists nothing against this paper. Absence here is not a guarantee, only a check that was made.

5 · Who and what money

Authors and funding

7 authors.

Mariana KarimovaDepartment of Biology and Biotechnologies Charles Darwin, The Sapienza University of Rome, 00185 Rome, Italy.ORCID 0000-0002-5348-0883
Giordana PonzianiDepartment of Biology and Biotechnologies Charles Darwin, The Sapienza University of Rome, 00185 Rome, Italy.
Diana VardanyanDepartment of Biology and Biotechnologies Charles Darwin, The Sapienza University of Rome, 00185 Rome, Italy.ORCID 0009-0009-3776-7639
Andrea AlfieriCentro Grandi Strumenti, University of Pavia, 27100 Pavia, Italy.ORCID 0000-0002-9716-0339
Maurizio ZuccottiDepartment of Biology and Biotechnology "Lazzaro Spallanzani", University of Pavia, 27100 Pavia, Italy.ORCID 0000-0003-4648-0663
Stefano TacconiDepartment of Biology and Biotechnologies Charles Darwin, The Sapienza University of Rome, 00185 Rome, Italy.ORCID 0000-0001-7424-1251
Luciana DiniDepartment of Biology and Biotechnologies Charles Darwin, The Sapienza University of Rome, 00185 Rome, Italy.ORCID 0000-0002-2633-9040

Funding

No grant is acknowledged in the PubMed record.

6 · The paper itself

Abstract

Extracellular vesicles (EVs) are attracting considerable interest due to their important role in cell signaling. However, their nanosized scale, complexity, and heterogeneity make even morphological characterization challenging. Only with the recent advances in cryogenic electron microscopy (cryo-EM), together with the increasing availability of cryo-electron microscopes, has it become possible to visualize the native structure of EVs. In this study we performed an in-depth cryo-EM analysis of EVs derived from four glioblastoma multiforme (GBM) cell lines (U87MG, U373MG, U251MG, and T98G), highlighting the morphological changes induced by temozolomide (TMZ) chemotherapeutic treatment. Size, shape, circularity, concentricity, membrane thickness, and electron density of EVs were analyzed. The key characteristic revealed by cryo-EM was that EVs can be enclosed not only by one membrane bilayer, but also by two or more bilayers (double-layered vesicles, DVs; and multilayered vesicles, MVs). Overall, TMZ treatment substantially modified both the morphology and production of EVs, decreasing the percentage of single-layered vesicles (SVs) while increasing that of DVs and MVs, as well as reducing the electron density of the EV cargo. Morphometric and morphological information can shed light on the contribution of EVs to tumor progression, metabolism, drug resistance, and immune evasion.

Indexed as

cryo-EMextracellular vesiclesglioblastoma multiformetemozolomide

Identifiers

PMID42274693
PMCPMC13257892

What Socratic holds

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Registered trials

None linked

Read under generation 80e0d062 · epoch 390. Bibliography from PubMed, PubMed Central and OpenAlex; grants from NIH RePORTER; trial links from ClinicalTrials.gov; estimates, votes and beliefs from the Socratic graph.